
Panthere de Diard
Neofelis diardi(G. Cuvier, 1823)
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Description
félin vivant à Bornéo et sur l'île de Sumatra
Source : Wikidata
Pays · région · aire protégée · écorégion · biome
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Calcul du tissu écologique de Neofelis diardi.
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Liste rouge IUCN
VU · Vulnérablecritères A2c; C1↘Décroissante- Évaluation
- 2015 · v3.1
- Altitude
- 0 – 1500 m
- Profondeur
- – m
État de la populationTexte officiel évaluation IUCNExpert
Our understanding of the Sunda Clouded Leopard’s population density on Borneo, and particularly in the Malaysian state of Sabah, has improved in recent years, with the advent of intensive, targeted camera traps surveys, analysed within a capture-recapture framework. Population density estimates from a range of forest types and elevations, and a gradient of disturbance are typically low (0.8 – 4.4 ind. / 100 km², with 6 of 10 surveys providing estimates of under 1.9 ind. / 100 km². All estimates from Sabah were derived using a spatially explicit capture recapture framework (SECR), from primary Dipterocarp forest, including lowland (Maliau Basin Conservation Area, 1.9 ind. / 100 km²: Brodie and Giordano 2012, Danum Valley Conservation Area, 1.76 ind. / 100 km²: Hearn, Ross and Macdonald unpublished data), upland (Tawau Hills National Park, 1.85 ind. / 100 km²: Hearn, Ross and Macdonald unpublished data) and submontane/montane (Crocker Range National Park, 1.4 ind. / 100 km²: Hearn, Ross and Macdonald unpublished data), population density estimates from selectively logged forests are more variable (Tangkulap-Pinangah Forest Reserve, 0.84 ind. / 100 km2, Segaliud Lokan Forest Reserve, 1.0 ind. / 100 km²: Wilting et al. 2012, Ulu Segama Forest Reserve, 2.55 ind. / 100 km², Tabin Wildlife Reserve, 2.26 ind. / 100 km² (Hearn, Ross and Macdonald unpublished data). In Kalimantan, there are only two studies that have yielded population density estimates of the Sunda Clouded Leopard. Loken pers. comm. used SECR and provide estimates from the selectively logged Wehea Forest in East Kalimantan (1.23 to 3.01 ind. / 100 km²). Using a conventional capture recapture framework, which typically results in higher densities from that derived using SECR, Cheyne et al. (2013) provide density estimates for a population of Sunda Clouded Leopards residing within the Sabangau peat swamp, a disturbed peat swamp forest in Central Kalimantan (0.7-4.4 ind. / 100 km²). Additional Sunda Clouded Leopard focused surveys on Borneo failed to find any evidence of this species, including the relatively small (ca. 43 km²) and isolated Kabili-Sepilok Forest Reserve, Sabah (Ross et al. 2013, Hearn, Ross and Macdonald unpublished data) and several forest sites in Kalimantan (Cheyne et al. in prep). Such surveys highlight the potential patchy nature of Sunda Clouded Leopard presence across the forested landscape of Borneo. The population density of this felid on Sumatra has received less attention, but surveys are currently underway (I. Haidir pers comm, 2015), and four camera surveys undertaken within the Kerinci landscape present a similar pattern to that of Borneo, with densities ranging from 0.8 to 1.6 individuals / 100 km² (Sollmann et al. 2014).
A Maximum Entropy based habitat suitability analysis for Borneo, discussed in detail above, provided a total AOO of 451,900 km² (Borneo: 378,900 km², Sumatra: 73,000 km²). Acknowledging that our estimate of current AOO may be biased high as a result of unmeasured hunting pressure, a potentially patchy distribution, and further loss of forest post-2010, we take a precautionary approach and estimate that the mean density of the Sunda Clouded Leopard across its AOO may plausibly be as low as 1 individual / 100 km². Extrapolation of this estimate to the wider AOO suggests that the number of mature individuals range-wide could be around 4,500 individuals, with around 3,800 persisting in Borneo and as few as 730 individuals remaining in Sumatra.
A Maximum Entropy based habitat suitability analysis for Borneo, discussed in detail above, provided a total AOO of 451,900 km² (Borneo: 378,900 km², Sumatra: 73,000 km²). Acknowledging that our estimate of current AOO may be biased high as a result of unmeasured hunting pressure, a potentially patchy distribution, and further loss of forest post-2010, we take a precautionary approach and estimate that the mean density of the Sunda Clouded Leopard across its AOO may plausibly be as low as 1 individual / 100 km². Extrapolation of this estimate to the wider AOO suggests that the number of mature individuals range-wide could be around 4,500 individuals, with around 3,800 persisting in Borneo and as few as 730 individuals remaining in Sumatra.
Menaces identifiées(9 menaces classées CMP-IUCN)
1_1Housing & urban areasOngoing2_1_1Shifting agricultureOngoing2_1_2Small-holder farmingOngoing2_1_3Agro-industry farmingOngoing2_2_3Scale Unknown/UnrecordedOngoing2_3_2Small-holder grazing, ranching or farmingOngoing5_1_1Intentional use (species is the target)Ongoing5_1_2Unintentional effects (species is not the target)Ongoing5_3_5Motivation Unknown/UnrecordedOngoing
Description complète des menacesTexte détaillé évaluation IUCNExpert
Habitat loss due to commercial logging and conversion to oil palm plantations and the increasing threat from poaching pose the greatest threats to the Sunda Clouded Leopard. Oil palm plantations are likely to expand in the future as a result of the push for biofuels, and forest cover on the island of Borneo, if current deforestation rates continue, is projected to decline from 50% to less than one-third by 2020 (Rautner 2005). A more recent analysis of deforestation risk predicts that by 2020 37% of the remaining forest in Borneo will have a greater than 25% risk of being deforested and that approximately 13% of Borneo’s forests (an area of 11,167 km²) will have a greater than 50% chance of being lost (Macdonald et al. in prep. a). This deforestation will not only reduce the area of available habitat for the Sunda Clouded Leopard but it is also likely to increase the fragmentation of its populations. In a modelling analysis of the likely impacts of this deforestation, Macdonald et al. (in prep. b) predict that between 2010 and 2020 the percentage of the landscape connected by dispersal in Borneo will decrease by over 60% and that this could lead to both a large reduction in the Sunda Clouded Leopard numbers and also a significant reduction in the potential gene flow between populations. Increased fragmentation of Clouded Leopard habitat may also be resulting in an increased risk of disease in some populations due to exposure to pathogens present in feral cat and dog populations. One study in Sabah found seropositive findings for canine parvovirus and/or distemper virus in feral cats and dogs, Malay Civets (Viverra tangalunga) and Common Palm Civets (Paradoxurus hermaphroditus) living sympatrically with Sunda Clouded Leopards (Nájera et al. in prep).
Poaching, either directly through the use of snares, or through the reduction of prey availability by the poaching of game species, poses a significant threat, both in Sumatra (Holden et al. 2001) and Borneo (Rabinowitz et al. 1987, Wilting et al. 2006, Cheyne et al. 2013).
Poaching, either directly through the use of snares, or through the reduction of prey availability by the poaching of game species, poses a significant threat, both in Sumatra (Holden et al. 2001) and Borneo (Rabinowitz et al. 1987, Wilting et al. 2006, Cheyne et al. 2013).
Habitats préférentiels (classification IUCN)
1_6Forest - Subtropical/Tropical Moist Lowland★1_9Forest - Subtropical/Tropical Moist Montane★1_7Forest - Subtropical/Tropical Mangrove Vegetation Above High Tide Level1_8Forest - Subtropical/Tropical Swamp
Mesures de conservation recommandéesStratégies de conservation IUCNExpert
The Sunda Clouded Leopard is currently listed under CITES Appendix I (as Neofelis nebulosa), and warrants full listing under N. diardi. The species is fully protected in Sumatra and Kalimantan (Indonesia), Sabah and Sarawak (Malaysia), and Brunei. It occurs in most protected areas along the Sumatran mountain spine, and in most of the larger protected areas on Borneo, though maintaining or re-establishing connectivity among protected areas (e.g. via habitat corridors) remains a critical conservation priority for the species. The Sunda Clouded Leopard has been the subject of increased research in recent years, but it remains one of the least studied of the world’s Pantherine felids, hampering the development of conservation actions. It is essential, therefore, to gain an understanding of this species’ distribution, abundance, and responses to anthropogenic modification, particularly in Sumatra, and also in Kalimantan, Sarawak and Brunei. Data on the distribution and abundance of this species in Sabah has improved greatly in recent years, and it remains essential to monitor the population over time. A detailed study of the Sunda Clouded Leopard’s basic ecology, including its diet and dispersal abilities is currently underway in Sabah. The threat posed by illegal hunting of this species, and the potential trade routes remain unclear, and should be the focus of research throughout its range.
Actions de conservation (11)Conservation Actions Classification Scheme — IUCNExpert
1_1Site/area protection2_1Site/area management3_1_1Harvest management3_1_2Trade management3_4_1Captive breeding/artificial propagation3_4_2Genome resource bank4_2Training4_3Awareness & communications5_4_1International level5_4_2National level5_4_3Sub-national level
Stress écologiques (15)Stresses Classification — IUCNExpert
1_1Ecosystem conversion1_1Ecosystem conversion1_1Ecosystem conversion1_1Ecosystem conversion1_1Ecosystem conversion1_1Ecosystem conversion1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation2_1Species mortality2_1Species mortality
Usage & commerce (3)Use & Trade — IUCNExpert
10Wearing apparel, accessoriesinternationalnationalsubsistance11Other household goodsinternationalnationalsubsistance16Establishing ex-situ production *
Priorités de recherche (8)Research Needed Classification — IUCNExpert
1_1Taxonomy1_2Population size, distribution & trends1_3Life history & ecology1_4Harvest, use & livelihoods1_5Threats1_6Actions2_1Species Action/Recovery Plan3_1Population trends
Niche IUCN globaleRealms · Systems · LMEs · Growth forms · FAOs — biogéographie IUCNExpert
Royaumes biogéographiques
Indomalayan
Systèmes (terrestre/eau douce/marin)
Terrestrial
Références bibliographiques (30)Sources scientifiques de l'évaluation IUCNExpert
- Macdonald, E.A., Cushman, S.A., Landguth, E.L., Hearn, A.J., Malhi, Y., and Macdonald, D.W. In prep B. Clouded leopard population connectivity and gene flow in the landscape: the past, present and future.
- Macdonald, E.A., Cushman, S.A., Landguth, E.L., Malhi, Y., and Macdonald, D.W. In prep A. Patterns and predictions of deforestation risk in Borneo.
- Nájera, F., Hearn, A.J., Evans, M.N., Guerrero-Sánchez, S., Nathan, S.K.S.S., de Gaspar, I., Revuelta, L. and Goossens, B. In prep. Serological survey for two canine pathogens in free-ranging and domestic carnivores in the Lower Kinabatangan Wildlife Sanctuary .
- IUCN. 2016. The IUCN Red List of Threatened Species. Version 2016-1. Available at: <a href="www.iucnredlist.org">www.iucnredlist.org</a>. (Accessed: 30 June 2016).
- McCarthy, J. L., Wibisono, H. T., McCarthy, K. P., Fuller, T. K., & Andayani, N. 2015. Assessing the distribution and habitat use of four felid species in Bukit Barisan Selatan National Park, Sumatra, Indonesia. . <i>Global Ecology and Conservation</i> 3: 210-221.
- Brodie, J. F., Giordano, A. J., Zipkin, E. F., Bernard, H., Mohd-Azlan, J. and Ambu. L. 2015. Correlation and persistence of hunting and logging impacts on tropical rainforest mammals. . <i>Conservation Biology </i> 29: 110-121.
- IUCN. 2015. The IUCN Red List of Threatened Species. Version 2015-4. Available at: <a href="www.iucnredlist.org">www.iucnredlist.org</a>. (Accessed: 19 November 2015).
- Mathai, J., Buckingham, L. and Ong, N. 2014. Borneo bay cat and other felids in a logging concession in Sarawak, Malaysian Borneo. <i>Cat News</i> 60: 34-35.
- Gaveau, D.L.A., Sloan, S., Molidena, E., Yaen, H., Sheil, D., Abram, N.K., Ancrenaz, M., Nasi, R., Quinones, M., Wielaard., N. and Meijaard, E. 2014. Four decades of forest persistence, clearance and logging on Borneo. <i>PLoS One</i> 9(7): e101654.
- Sollmann, R., Linkie, M., Haidir, I. A., & Macdonald, D. W. 2014. Bringing clarity to the clouded leopard Neofelis diardi: first density estimates from Sumatra. <i>Oryx</i> 48(4): 536-539.
- Pusparini, W., Wibisono, H.T., Reddy, G.V., Tarmizi, T. and Bharata, P. 2014. Small and medium sized cats in Gunung Leuser National Park, Sumatra, Indonesia. <i>Cat News</i> Special Issue(8): 4-9.
- Hearn, A. J., Ross, J., Pamin, D., Bernard, H., Hunter. L., & Macdonald, D. W. 2013. Insights into the spatial and temporal ecology of the Sunda clouded leopard Neofelis diardi. <i>Raffles Bulletin of Zoology</i> 61(2): 871-875.
- Pacifici, M., Santini, L., Di Marco, M., Baisero, D., Francucci, L., Grottolo Marasini, G., Visconti, P., Rondinini, C. 2013. Generation length for mammals. <i>Nature Conservation </i> 5: 87-94.
- Ross, J., Hearn, A. J., Johnson, P. J., & Macdonald, D. W. 2013. Activity patterns and temporal avoidance by prey in response to Sunda clouded leopard predation risk. . <i>Journal of Zoology</i> 290(2): 96-106.
- Nájera, F., Bolongon, G., Abram, N., Goossens, B., Ambu, L., Macdonald, D.W, and Hearn, A.J. 2013. Observation of a road-killed Sunda clouded leopard in Malaysian Borneo. . <i>Cat News</i> 58: 42-43.
- Cheyne, S. M., Stark, D. J., Limin, S. H., & Macdonald, D. W. 2013. First estimates of population ecology and threats to Sunda clouded leopards Neofelis diardi in a peat-swamp forest, Indonesia. <i>Endangered Species Research</i> 22: 1-9.
- Miettinen, J., Shi, C., Tan, W. J., & Liew, S. C. 2012. 2010 land cover map of insular Southeast Asia in 250-m spatial resolution. <i>Remote Sensing Letters</i> 3: 11-20.
- Brodie, J. and Giordano, A. J. 2012. Density of the Vulnerable Sunda clouded leopard Neofelis diardi in a protected area in Sabah, Malaysian Borneo. . <i>Oryx</i> 46: 427-430.
- Wilting, A., Mohamed, A., Ambu, L. N., Lagan, P., Mannan, S., Hofer, H., & Sollmann, R. 2012. Sunda Clouded Leopard Neofelis diardi density in two used forests in Sabah, Malaysian Borneo. . <i>Oryx</i> 46: 423-426.
- Bernard, H. Baking, E.L., Matsubayasi, M., and Ahmad, A.H. 2012. Records of Bornean felids in and around Tabin Wildlife Reserve, Sabah, Malaysia. . <i>Cat News</i> 56: 4-7.
- Wilting, A., Christiansen, P., Kitchener, A. C., Kemp, Y. J. M., Ambu, L., & Fickel, J. 2011. Geographical variation in and evolutionary history of the Sunda clouded leopard (Neofelis diardi) (Mammalia: Carnivora: Felidae) with the description of a new subspecies from Borneo. <i>Molecular Phylogenetics and Evolution</i> 58: 317-328.
- Cheyne, S.M. and Macdonald, D.W. 2011. Wild felid diversity and activity patterns in Sabangau peat-swamp forest, Indonesian Borneo. <i>Oryx</i> 45: 119-124.
- Morino, L. 2010. Clouded leopard predation on a wild juvenile siamang. <i>Folia Primatologica</i> 81: 362-368.
- Ross, J., Hearn, A. J., Bernard, H., Secoy, K., & Macdonald, D. W. 2010. A framework for a Wild Cat Action Plan for Sabah. Global Canopy Programme, Oxford, UK.
- Mohamed, A., Samejima, H., and Wilting, A. 2009. Records of five Bornean cat species from Deramakot Forest Reserve in Sabah, Malaysia. . <i>Cat News</i> 51: 14-17.
- Ridout, M. S., & Linkie, M. 2009. Estimating overlap of daily activity patterns from camera trap data. <i>Journal of Agricultural, Biological, and Environmental Statistics</i> 14(3): 322-337.
- Christiansen, P. 2009. Neotype of Neofelis diardi (Mammalia: Carnivora: Felidae). . <i>Zootaxa </i> 2110: 58-68.
- Matsuda, I., Tuuga, A., & Higashi, S. 2008. Clouded leopard (Neofelis diardi) predation on proboscis monkeys (Nasalis larvatus) in Sabah, Malaysia. <i>Primates</i> 49: 227-231.
- Christiansen, P. 2008. Species distinction and evolutionary differences in the clouded leopard (Neofelis nebulosa) and Diard’s clouded leopard (Neofelis diardi). . <i>Journal of Mammology</i> 89: 1435–1446.
- Gordon, C. H. and Stewart, A.-M. E. 2007. The use of logging roads by clouded leopards. <i>Cat News</i> 47: 12-13.
Évaluateurs & contributeurs (3)Personnes ayant contribué à l'évaluation IUCNExpert
assessor
Hearn, A., Ross, J., Brodie, J., Cheyne, S., Haidir, I.A., Loken, B., Mathai, J., Wilting, A. & McCarthy, J.
contributor
Sanderson, J. & Sunarto, S.
evaluator
Nowell, K., Hunter, L., Duckworth, J.W., Breitenmoser-Würsten, C., Lanz, T. & Breitenmoser, U.
⚠ 1 erratum publié après l'évaluation.
Hearn, A., Ross, J., Brodie, J., Cheyne, S., Haidir, I.A., Loken, B., Mathai, J., Wilting, A. & McCarthy, J. 2015. Neofelis diardi (errata version published in 2016). The IUCN Red List of Threatened Species 2015: e.T136603A97212874. Accessed on 05 May 2026.
Distribution mondiale
Calcul de la distribution GBIF· ~10–60 s
Phénologie
Calcul du calendrier d'apparition· ~5–30 s
Consulter sur les bases externes
Observations & statuts
Cartographie
Note nomenclaturale & synonymesDétails taxonomiques + synonymes CoLExpert
Note nomenclaturale
TAXREF v18 — INPN/MNHNSynonymes (2)— redirigent vers cette page
- Felis diardiG. Cuvier, 1823
- Neofelis nebulosa diardi(G. Cuvier, 1823)
Sources : Catalogue of Life Cross-References (synonymes) · TAXREF v18 INPN/MNHN (commentaires FR).