
Palmier de Bankoualé
Livistona carinensis(Chiov.) J.Dransf. & N.W.Uhl
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Calcul du tissu écologique de Livistona carinensis.
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Liste rouge IUCN
EN · En dangercritères A2c; B2ab(iii,v)↘Décroissante- Évaluation
- 2018 · v3.1
- Altitude
- 165 – 970 m
- Profondeur
- – m
État de la populationTexte officiel évaluation IUCNExpert
In Djibouti, the majority of trees are over-mature, and the majority of the seeding trees are expected to die from old age over a short period of time. The removal of young palms in gardens, coupled with the livestock-grazing and flash-flooding events outside gardens, greatly limit the recruitment of young palms (Welch and Welch 1999, Ford et al. 2008). This is likely to leave very small, fragmented subpopulations at 21 small sites.
In Somalia, juvenile palms (2-10 m of trunk, sexually mature) have never been reported. However, hundreds of young rosette palms (less than 2 m of trunk, not sexually mature) were recorded at each of the five sites in 1983-1985 (C. Barbier pers. comm. 1987), mostly growing along the irrigation ducts within the date palm plantations. The last observations of the species in Somalia dated from 1995 and 1997 when 11 and 27 mature palms (trunks of 10-40 m, sexually mature) were respectively counted at Galgala and Tisjiec (M. Thulin pers. comm. 1998).
By contrast, in Yemen, especially at Al Jubah, most subpopulations are comprised of bushy young plants (rosettes and juveniles trees), with very few tall old palms, and are thus presumed to be reproductively healthy. Indeed, Al Jubah appears to hold the largest and most intact stands of the Bankoualé Palm. Hubaishan and Bahah (M.A. Hubaishan pers. comm. 2004), estimated the number of mature individuals at 2,000 (adults and juveniles palms). Observing Google Earth 2016 satellite imagery, several dense stands can be seen, with a clumped distribution along the wadi for a distance of 5-6 km. The stands at Al-Ghyadhat and Wadi Al Himae are isolated and small, respectively one and ca. 90 mature individuals counted in 2003/2004 (M.A. Hubaishan pers. comm. 2004, Kilian et al. 2004).
The population trend for the species in Dijbouti can be inferred from subpopulation counts made between 1985 and 2015 (Welch and Welch 1999, Ford and Bealy 2004, Ford et al. 2008, A. Laurent pers. comm. 2015). The data which include the most sites (18) shows a decline of 21.7% over eight years. A longer set of data (12 sites), reveals a decline of 50% over 21 years (1985-2006). For the four largest sites (accounting for ca. 50% of the entire subpopulation for the country in 1985) there is data for the longest time period, and this shows a decline of 75.5% over 30 years. From these observed data, a population decline of at least 50% is inferred over three generations (60 years). At only one site (Satabou south) has a population increase been recorded, an increase of 23 adults (135%) over 10 years.
In Somalia, the number of mature individuals declined by 76% in 23 years, from 158 in 1973 (C. Barbier pers. comm. 1987) to 38 in 1997 (M. Thulin pers. comm. 1998). Due to the maturity of all recorded individuals and the lack of reports of young plants or seedlings when surveyed in 1997, it is not unreasonable to assume that the species is now at least functionally extinct in this region and the subpopulations in Somalia have experienced a decline of 100% over two generations (40 years). By extrapolation, it is inferred that they have experienced a decline of 100% over three generations (60 years).
In Yemen, mature palms were counted at Al Ayn and Al Mintaq in 1998 (Welch and Welch 1998) and 2004 (M.A. Hubaishan pers. comm. 2004). An 80% decline was estimated to have taken place over the six years, with the number of mature individuals falling from 970 to 196. In 1998 a total of 1936 stumps were also recorded - suggesting a total earlier population size of 2906, and indicating a total loss of at least 93%. Despite this, examination of 2016 satellite imagery (Google Earth), reveals many Bankoualé palms still to be present, though they are scattered among the more numerous date palms. There is not sufficient data on all the Yemen sub-populations to make an overall estimate of the degree of continuing decline over three generations. However, whilst the Al Jubah subpopulation at ca. 2,000 mature individuals is the most important, it can be inferred that the subpopulations in Yemen have overall experienced a continuing decline from 1989 to today.
Considering the best available data, a continuing decline in subpopulation sizes, over three generations, is inferred to be at least 50% in Djibouti, 100% in Somalia and occurring but at an undetermined rate in Yemen, leading to an inferred global continuing decline in the global population size of the species of at least 50%. The current global population for the species is estimated to comprise 2,580 mature individuals (ca. 2,300 in Yemen and 280 in Djibouti).
Menaces identifiées(10 menaces classées CMP-IUCN)
2_1_2Small-holder farmingRapid DeclinesMajority (50-90%)Ongoing2_2_1Small-holder plantationsRapid DeclinesMajority (50-90%)Ongoing2_3_2Small-holder grazing, ranching or farmingRapid DeclinesMinority (<50%)Ongoing5_3_1Intentional use: (subsistence/small scale) [harvest]Rapid DeclinesMajority (50-90%)Ongoing11_2DroughtsSlow, Significant DeclinesMinority (<50%)Ongoing11_4Storms & floodingSlow, Significant DeclinesMinority (<50%)Ongoing12_1Other threatSlow, Significant DeclinesMinority (<50%)Ongoing4_1Roads & railroadsSlow, Significant DeclinesMajority (50-90%)Ongoing5_3_3Unintentional effects: (subsistence/small scale) [harvest]Slow, Significant DeclinesMinority (<50%)Ongoing7_2_3Abstraction of surface water (agricultural use)Slow, Significant DeclinesMajority (50-90%)Ongoing
Description complète des menacesTexte détaillé évaluation IUCNExpert
In Somalia, cutting the palm for building (because the palm trunks are the only local long straight timber available) and the establishment and expansion of date palm plantations appears to have led to the disappearance of the palm in this country.
In Yemen, the major threats are also cutting for building timber and the establishment and expansion of date palm plantations (Welch and Welch 1998, M.A. Hubaishan pers. comm. 2004). However, here, perhaps because the stands and areas of suitable habitat are large, some mature young palms continue to grow among the date palm plantations at Al Ayn and Al Mintaq. The Al Jubah stands still appears intact, but it can be inferred that an increase in the unsustainable cutting of trunks for timber and expansion of the date palm plantations is likely to have occurred following the opening of an access track for vehicles in August 2004. Only the mature palms are cut because, as in Somalia, they are the only local long straight timber and are thus a valuable building material. Piles of trunks cut and ready for building use were observed at Al Mintaq in 1989 (Bazara et al. 1990) and 1998 (Welch and Welch 1998) and at Al Jubah in 1998 (Welch and Welch 1998) and 2004 (M.A. Hubaishan pers. comm. 2004).
In Djibouti, the major threats to the Bankoualé Palm are habitat loss due to agricultural activities, habitat degradation due to diversion of surface water to irrigate gardens, torrential floods and livestock grazing (Welch and Welch 1999, Ford et al. 2008). The first rural gardens were established for the production of mangoes when cheap labour was provided by the arrival of Ethiopian immigrants in the 1960s. The gardens have been built on terraces along the wadi sides in the same places where the palm grows. Although adult and small rosette palms are tolerated, bushy young palms are generally removed to make space for cultivation, and young plants are grazed by livestock. Moreover, in order to develop gardens the surface water is channelled for irrigation: as a result the only damp areas, which are protected from flash-floods and where the palm can germinate and become established, are in the gardens, putting the gardens and the palm in direct conflict. Outside the gardens, the suitable habitat for germination and establishment of young individuals has become rare due to the diversion of surface water for irrigation. The young plants are grazed by livestock, greatly reducing the survival of young plants. Additionally, the natural and regular flash-flooding events, wash away huge number of seedlings (Ford et al. 2008). These flash-floods, have become increasingly severe due to the gradual denuding of the water catchment. This started with the localized cutting of Buxus hildebrandtii bushes on the wadi sides in the early 1970s (Ethiopian immigrants used them for thatching their traditional houses) and, in 2014 became a slow but systematic removal of timber from the whole Goda massif water catchment, driven by the growing and lucrative demand for charcoal in the capital (A. Laurent pers. comm. 2017). It is also reported that adult palms are progressively uprooted due to violent summer winds (A. Laurent pers comm. 2017). These major threats are exacerbated by a drought in the region since 2009 which has led to the drying up of permanent springs and profound changes to both the wadi bed habitats used by the palm and the hydrological regime (A. Laurent pers. comm. 2017).
Habitats préférentiels (classification IUCN)
14_4Artificial/Terrestrial - Rural Gardens★5_9Wetlands (inland) - Freshwater Springs and Oases★8_1Desert - Hot★14_3Artificial/Terrestrial - Plantations15_7Artificial/Aquatic - Irrigated Land (includes irrigation channels)
Mesures de conservation recommandéesStratégies de conservation IUCNExpert
Livistona carinensis does not occur in any effectively protected areas and there is no coordinated international work on the species. The species is grown ex situ in botanic gardens (BGCI, 2017), but populations in cultivation are small and not yet self-supporting. There have been successful ex situ germination trials in Yemen (S.O. Bahah pers. comm. 2016) but this work has been disrupted by civil war. Seeds from Djibouti collected in August 2007 have been deposited at the Millennium Seed Bank (UK).
Community-based conservation areas and/or micro reserves are needed to provide a suitable and practical level of protection for the remaining stands of the palm. As the largest remaining stand of L. carinensis in the world, Al Jubah in Yemen should be a priority site for in situ conservation. Effective restrictions on unsustainable harvesting for timber are needed to ensure the continued survival of the palm in Yemen. Shapcott et al. (2009) indicated that the subpopulations in Yemen (and Somalia if they still exist), are highly significant for the conservation of the genetic diversity of the palm.
In Djibouti, there have been positive signs of local conservation initiatives. A few young palms have been allowed to grow in peripheral areas in two gardens at Bankoualé, and some seed has been distributed in Tadjourah and Djibouti city to encourage horticultural interest in growing the palm in private gardens (A. Laurent pers. comm. 2017). At a national level, Djibouti’s 2017 National Biodiversity Strategy and Action Plan (Direction de l'Environnement et du Développement Durable 2017) identifies replenishment of the underground aquifers as the primary issue for both the continuation of agricultural activities and the palm at Bankoualé, and recommends that establishment of “biodiversity conservation areas” in private gardens be integrated as much as possible into watershed management. The restoration of the Buxus on the mountainsides above the main wadi systems in Djibouti would help to reduce the incidence and severity of flash-floods and their impact on palm seedling survival.
In Somalia, searches should be initiated to look for any remaining palm stands, using Species Distribution Modelling and/or study of satellite imagery to pinpoint potential unexplored sites of suitable habitat where the species might be able to grow. It would also be worth carrying out feasibility studies for the re-introduction of the palm in Somalia, if suitable sites can be identified.
Actions de conservation (8)Conservation Actions Classification Scheme — IUCNExpert
1_1Site/area protection2_1Site/area management2_3Habitat & natural process restoration3_1_1Harvest management3_3_1Reintroduction3_4_1Captive breeding/artificial propagation3_4_2Genome resource bank4_3Awareness & communications
Stress écologiques (27)Stresses Classification — IUCNExpert
1_1Ecosystem conversion1_1Ecosystem conversion1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation1_2Ecosystem degradation1_3Indirect ecosystem effects1_3Indirect ecosystem effects1_3Indirect ecosystem effects1_3Indirect ecosystem effects2_1Species mortality2_1Species mortality2_1Species mortality2_1Species mortality2_1Species mortality2_1Species mortality2_1Species mortality2_1Species mortality2_2Species disturbance2_2Species disturbance2_2Species disturbance2_2Species disturbance2_3_2Competition2_3_2Competition2_3_7Reduced reproductive success
Usage & commerce (4)Use & Trade — IUCNExpert
1Food - humansubsistance12Handicrafts, jewellery, etc.subsistance7Fuelssubsistance9Construction or structural materialssubsistance
Priorités de recherche (5)Research Needed Classification — IUCNExpert
1_2Population size, distribution & trends2_1Species Action/Recovery Plan2_2Area-based Management Plan3_1Population trends3_2Harvest level trends
Niche IUCN globaleRealms · Systems · LMEs · Growth forms · FAOs — biogéographie IUCNExpert
Royaumes biogéographiques
Systèmes (terrestre/eau douce/marin)
Formes de croissance
Références bibliographiques (22)Sources scientifiques de l'évaluation IUCNExpert
- IUCN. 2018. The IUCN Red List of Threatened Species. Version 2018-1. Available at: <a href="www.iucnredlist.org">www.iucnredlist.org</a>. (Accessed: 28 June 2018).
- Direction de l'Environnement et du Développement Durable. 2017. Stratégie et programme d'action natinaux pour la biodiversité. Ministère de l'Habitat, de l'Urbanisme et de l'Environnement.
- Google Earth. 2016. <i>Satellite Imagery</i>.
- Bacon, C., Baker, W.J. and Simmons M.P. 2012. Miocene dispersal drives island radiations in the palm Tribe Trachycarpeae (Arecaceae). <i>Systematic Biology</i> 61(3): 426-442.
- Bachman, S., Moat, J., Hill, A.W., de la Torre, J. and Scott, B. 2011. Supporting Red List threat assessments with GeoCAT: geospatial conservation assessment tool. In: V. Smith and L. Penev (eds) e-Infrastructures for data publishing in biodiversity science. <i>Zookeys</i> 150: 117–126.
- Shapcott, A., Dowe, J.L. and Ford, H. 2009. Low genetic diversity and recovery implications of the vunerable Bankoualé Palm <i> Livistona carinensis </i> (Arecaceae), from north-eastern Africa and the southern Arabian Peninsula. <i>Conservation Genetics</i> 10: 317-327.
- Dowe, J.L. 2009. A taxonomic account of <i>Livistona</i> R.Br. (Arecaceae). <i>Gardens Bulletin Singapore</i> 60(2): 185-344.
- Ford, H., Conochie, G., Trathan, P. and Gillet, H. 2008. The decline of the Bankoualé Palm in Djibouti: El Nino and changes in architectural fashion. <i>Palms</i> 52(2): 89-95.
- Ford, H. and Bealy, C. 2004. Status of the Bankoualé Palm, <i>Livistona carinensis</i>, in Djibouti. <i>Palms</i> 48(2).
- Kilian, N., Hein, P. and Hubaishan, M.A. 2004. Further notes on the flora of the southern coastal mountains of Yemen. <i>Willdenowia</i> 34: 159-182.
- El Mashjary, M.S., Hein, P. and Killian, N. 2001. The Endangered Fan Palm <i>Livistona carinensis</i> in Yemen. <i>Yemeni Journal of Science</i> 3(1): 21-25.
- Welch, H. and Welch, G. 1999. A report on the birds of Djibouti and the Bankoualé Palm <i>Livistona carinensis</i>. Ministère de l'Environnement, du Tourisme et de l'Artisanat. Direction de l'Environnement, Djibouti.
- Welch, H and Welch, G. 1998. Survey of the Bankoualé Palm <i>Livistona carinensis</i>.
- Bazara'a, M., Guarino, L., Miller, A. and Obadi, N. 1990. Observations of an endangered palm in Arabia. <i>Edinburgh Journal of Botany</i> 47(3): 375-379.
- Barbier, C. 1985. Further notes on <i>Livistona carinensis</i> in Somalia. <i>Principes</i> 29(4): 151-155.
- Dransfield, J. and Uhl, N.W. 1983. <i>Wissmannia</i> (Palmae) </i>Reduced <i>Livistona</i>. <i>Kew Bulletin</i> 38(2): 199-200.
- Chédeville, E. 1972. La végétation du Territoire Français des Afars et des Issas. <i>Webbia</i> 26(2): 243-266.
- Tomlinson, P.B. 1961. The problem of <i>Wissmannia</i>. <i>Principes</i> 5: 33-34.
- Monod, T. 1955. Remarques sur un palmier peu connu: <i>Wissmannia carinensis</i> (Chiov) Burret. <i>Bulletin de l'Institut Fondamental d'Afrique Noire</i> 17: 338-358.
- Burret, M. 1943. Die Palmen Arabiens. <i>Botanische Jahrbücher für Systematik, Planzengeschichte und Planzengeographie</i> 73: 175-190.
- Aubert de la Rüe, E. 1939. <i>La Somalie Française</i>. Librairie Gallimard, Paris.
- Chiovenda, E. 1929. <i>Flora Somala</i>. Sind. Ital. Arti Grafiche, Rome.
Évaluateurs & contributeurs (4)Personnes ayant contribué à l'évaluation IUCNExpert
Cosiaux, A., Welch, H., Gardiner, L.M., Welch, G. & Couvreur, T.L.P. 2018. Livistona carinensis. The IUCN Red List of Threatened Species 2018: e.T30402A95306943. Accessed on 05 May 2026.
Distribution mondiale
Phénologie
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Observations & statuts
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Note nomenclaturale & synonymesDétails taxonomiques + synonymes CoLExpert
Note nomenclaturale
TAXREF v18 — INPN/MNHNSynonymes (2)— redirigent vers cette page
- Hyphaene carinensisChiov.
- Wissmannia carinensis(Chiov.) Burret
Sources : Catalogue of Life Cross-References (synonymes) · TAXREF v18 INPN/MNHN (commentaires FR).